Cornus

Cornus Temporal range: Late Campanian–Holocene, 73–0 Ma[1] PreꞒ Ꞓ O S D C P T J K Pg N
Cornus kousa var. chinensis
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Clade:	Angiosperms
Clade:	Eudicots
Clade:	Asterids
Order:	Cornales
Family:	Cornaceae
Genus:	Cornus L.
Type species
Cornus mas L.
Subgenera
Afrocrania Arctocrania Cornus Cynoxylon Discocrania Kraniopsis Mesomora Sinocornus Syncarpea Yinquania
Synonyms
Chamaepericlimenum Hill

Cornus is a genus of about 30–60 species^{[Note 1]} of woody plants in the family Cornaceae, commonly known as dogwoods or cornels, which can generally be distinguished by their blossoms, berries, and distinctive bark.^[3] Most are deciduous trees or shrubs, but a few species are nearly herbaceous perennial subshrubs, and some species are evergreen. Several species have small heads of inconspicuous flowers surrounded by an involucre of large, typically white petal-like bracts, while others have more open clusters of petal-bearing flowers. The various species of dogwood are native throughout much of temperate and boreal Eurasia and North America, with China, Japan, and the southeastern United States being particularly rich in native species. The genus is also known from South America with members such as Cornus peruviana.

Species include the common dogwood Cornus sanguinea of Eurasia, the widely cultivated flowering dogwood (Cornus florida) of eastern North America, the Pacific dogwood Cornus nuttallii of western North America, the Kousa dogwood Cornus kousa of eastern Asia, and two low-growing boreal species, the Canadian and Eurasian dwarf cornels (or bunchberries), Cornus canadensis and Cornus suecica respectively.

Depending on botanical interpretation, the dogwoods are variously divided into one to nine genera or subgenera; a broadly inclusive genus Cornus is accepted here.

Terminology

Cornus is the Latin word for the cornel tree, Cornus mas. The name cornel dates to the 1550s, via German from Middle Latin cornolium, ultimately from the diminutive cornuculum, of cornum, the Latin word for the cornel cherry. Cornus means "horn",^[4] presumably applied to the cherry after the example of κερασός, the Greek word for "cherry", which itself is of pre-Greek origin but reminiscent of κέρας, the Greek word for "horn".

The name "dog-tree" entered the English vocabulary before 1548, becoming "dogwood" by 1614. Once the name dogwood was affixed to this kind of tree, it soon acquired a secondary name as the hound's tree, while the fruits came to be known as "dogberries" or "houndberries" (the latter a name also for the berries of black nightshade, alluding to Hecate's hounds).^{[citation needed]}

The name was explained, from as early as the 16th century itself, as derived from dag "skewer",^[5][6] as the wood of the tree was said to have been used to make butcher's skewers. This is uncertain, as the form *dagwood was never attested. It is also possible that the tree was named for its berry, called dogberry from at least the 1550s, where the implication could be that the quality of the berry is inferior, as it were "fit for a dog".^[7]

An older name of the dogwood in English is whipple-tree, occurring in a list of trees (as whipultre) in Geoffrey Chaucer Canterbury Tales.^[8] This name is cognate with the Middle Low German wipel-bom "cornel", Dutch wepe, weype "cornel" (the wh- in Chaucer is unetymological, the word would have been Middle English wipel). The tree was so named for waving its branches, cf. Middle Dutch wepelen "totter, waver", Frisian wepeln, German wippen.^[9]

The name whippletree, also whiffle-tree, now refers to an element of the traction of a horse-drawn cart linking the draw pole of the cart to the harnesses of the horses in file. In this sense it is first recorded in 1733. This mechanism was usually made from oak or ash (and not from dogwood), and it is unlikely that there is a connection to the name for whipple-tree for Cornus.^[10]

Description

Dogwoods have simple, untoothed leaves with the veins curving distinctively as they approach the leaf margins. Most dogwood species have opposite leaves, while a few, such as Cornus alternifolia and C. controversa, have their leaves alternate. Dogwood flowers have four parts. In many species, the flowers are borne separately in open (but often dense) clusters, while in various other species (such as the flowering dogwood), the flowers themselves are tightly clustered, lacking showy petals, but surrounded by four to six large, typically white petal-like bracts.^{[citation needed]}

The fruits of all dogwood species are drupes with one or two seeds, often brightly colorful. The drupes of species in the subgenus Cornus are edible. Many are without much flavor. Cornus kousa and Cornus mas are sold commercially as edible fruit trees. The fruits of Cornus kousa have a sweet, tropical pudding like flavor in addition to hard pits. The fruits of Cornus mas are both tart and sweet when completely ripe. They have been eaten in Eastern Europe for centuries, both as food and medicine to fight colds and flus. They are very high in vitamin C. By contrast, the fruits of species in subgenus Swida are mildly toxic to people, though readily eaten by birds.^{[citation needed]}

Dogwoods are used as food plants by the larvae of some species of butterflies and moths, including the emperor moth, the engrailed, the small angle shades, and the following case-bearers of the genus Coleophora: C. ahenella, C. salicivorella (recorded on Cornus canadensis), C. albiantennaella, C. cornella and C. cornivorella, with the latter three all feeding exclusively on Cornus.^{[citation needed]}

Uses

Dogwoods are widely planted horticulturally, and the dense wood of the larger-stemmed species is valued for certain specialized purposes: cutting boards and fine turnings can be made from the fine-grained wood. Over 32 different varieties of game birds, including quail, feed on the dogwoods’ red seeds.^[11]

Classification

The following classification recognizes a single, inclusive genus Cornus,^[24][25] with four subgroups and ten subgenera supported by molecular phylogeny.^[26][27][28] Geographical ranges as native plants are given below.

Cultural references

The inflorescence of the Pacific dogwood (Cornus nuttallii) is the official flower of the province of British Columbia. The flowering dogwood (Cornus florida) and its inflorescence are the state tree and the state flower respectively for the U.S. Commonwealth of Virginia. It is also the state tree of Missouri and the state flower of North Carolina,^[35] and the state memorial tree of New Jersey.^[36] The term "dogwood winter", in colloquial use in the American Southeast, especially Appalachia,^[37] is sometimes used to describe a cold snap in spring, presumably because farmers believed it was not safe to plant their crops until after the dogwoods blossomed.^[38]

Notes

References

External links

Wikimedia Commons has media related to Cornus.

Wikispecies has information related to Cornus.

• Dogwood history and uses Archived 2016-03-03 at the Wayback Machine
• Cornus in Flora of China